The main changes of the inner structure of some spiriferid genera from Paleothetis in Carboniferous time were investigated: these were an appearance of the cross fi ssures on the area of pedical valve; appearance and evolution of the vascular system; reduction of the fi eld of ovarian pits and evolution of the tooth plates and other. On that base the assumption about hypothetical philogenesis of the some Carboniferous spiriferid genera was done and the general borders where the Carboniferous spiriferid system was changed were observed.

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